Holland Taylor : Peggy Peabody

Helena's eccentric millionaire mother, has been a regular recurring character since season one.

Dallas Roberts : Angus Partridge

Angelica's manny pursued Kit and finally managed to win her over.

Jane Lynch : Joyce Wischnia

Unscrupulous lawyer, Joyce first tried making a pass at Tina and then represented Bette in trying to gain sole custody of Angelica.

Meredith McGeachie : Tonya

Love or hate her, you can't help but enjoy any scene with Tonya, Dana's rather overbearing fiancee.

Alexandra Hedison : Dylan Moreland

Helena had finally thought she had found true love, but Dylan wasn't the innocent straight girl she seemed.

Alan Cumming : Billie Blaikie

Billy came along to help Kit manage the Planet and advise Moria on how to become Max.

Rosanna Arquette : Cherie Jaffee

Charie Jaffe turned the tables on player Shane in season one and returned in seasone three to mess up her life a little more.

Eric Roberts : Gabriel McCutcheon

When Shane met her dad, he turned out to be what she always thought she was.

Sandra Bernhard : Charlotte Birch

Charlotte called Jenny on her poor writing and then took her under her wing.

Ossie Davies : Melvin Porter

Kit and Bette's judgemental father, passed away at the end of season 2. Ossie Davies himself passed away before season 2 aired.

Camryn Manheim : Veronica Bloom

obnoxious producer that took a liking to Shane.

Kelly Lynch : Ivan Aycock

the drag king tried to woo Kit, but turned out to be another player.

Anne Ramsey : Robin

Robin fell for Jenny, but came on a little too strong.

Anne Archer : Lenore Pieszecki

excellent as Alice's flaky mother Lenore

Ion Overman : Candace Jewel

the infamous carpenter that caught Bette's eye and sparked of a series of events that led to the downfall of tibette.

Julian Sands : Nick Barashkov

a one episode appearance as Jenny's old tutor, was memorable for his speech on lies.

Snoop Dogg : Slim Daddy

a guest appearance as a rap artist interested in mixing one of Kit's songs.